Principal Engineer, AI Security

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Principal Engineer, AI Security in USA

Lila Sciences
Full time
Email

USA
10/01/2026
Report

Cambridge, MA

Responsibilities

Translate the AI security roadmap into delivered controls with measurable outcomes.
Design and implement security controls for training and inference pipelines, model registries, MCP servers, and agent runtimes.
Maintain continuous compliance of AI infrastructure against internal standards and applicable frameworks.
Drive vulnerability remediation across AI infrastructure, models, and dependencies through closure with system owners.
Reduce risk across data, models, pipelines, and agentic systems through threat modeling, control design, and adversarial assessment.
Partner with AI Platform, AI Research, ML Operations, Science, and AI Safety to embed security at design time.

Requirements

Strong background in security engineering, cloud security, or DevSecOps with hands-on experience securing production infrastructure.
Working knowledge of AI/ML systems, the ML lifecycle, and AI-specific risks including prompt injection, model and data poisoning, insecure agent tooling, and inference-time threats.
Experience with software supply chain security and dependency and vulnerability management.
Experience implementing cloud data security controls including classification, data flow mapping, access control, and encryption.
Proven ability to partner across engineering teams and drive cross-functional remediation to closure.
Comfort operating with ambiguity in a fast-moving research environment.
Bonus: hands-on AWS security experience, SBOM tooling at scale, familiarity with AI governance and assurance frameworks such as NIST AI RMF, or MAESTRO-style threat modeling for agentic systems.

Benefits

Full-time U.S. employees receive medical, dental, and vision coverage; employer-paid life and disability insurance; flexible time off; generous company-wide holidays; paid parental leave; educational assistance; commuter benefits including bike share memberships for office-based employees; and a subsidized lunch program.
Full-time employees outside the U.S. receive a comprehensive benefits program tailored to their region.
The role offers bonus potential and generous early-stage equity.
The position is full-time; work arrangement and contract duration are not stated.

Salary: $252k - $374k/yr

Lila Sciences

Lila Sciences is building the world's first scientific superintelligence platform and autonomous laboratories tailored for life, chemical, and materials sciences. Our innovative approach applies AI to every aspect of the scientific method, enabling researchers to tackle significant global challen...

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